Nobel Prize in Physiology or Medicine

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Emil Adolf von Behring was the first person to receive the Nobel Prize in physiology or Medicine, for his work on the treatment of diphtheria.
Emil Adolf von Behring was the first person to receive the Nobel Prize in physiology or Medicine, for his work on the treatment of diphtheria.

Below is a list of Nobel Prize laureates in Physiology or Medicine from 1901 to the present.[1]

The state of some countries changed from when a Nobel laureate lived in the country, became a laureate, and today. The flag that precedes the country of citizenship reflects the earlier of the following two situations: (1) the state of the country when the laureate emigrated or (2) the state of the country when he/she became a Nobel laureate. Some laureates obtained additional citizenships in their lifetime. For those with multiple citizenships, the year they obtained an additional citizenship, when known, precedes the country of citizenship. For laureates who obtained additional citizenships after winning the Nobel prize, the flag that precedes the country of citizenship reflects the earlier of the following two situations: (1) the state of the country at their death or (2) the current state of the country.

Year Laureate, Country of Citizenship Reason
1901 Emil Adolf von Behring, Flag of German Empire German Empire "for his work on serum therapy, especially its application against diphtheria, by which he has opened a new road in the domain of medical science and thereby placed in the hands of the physician a victorious weapon against illness and deaths"[2]
1902 Ronald Ross, Flag of the United Kingdom United Kingdom "for his work on malaria, by which he has shown how it enters the organism and thereby has laid the foundation for successful research on this disease and methods of combating it"[3]
1903 Niels Ryberg Finsen, Flag of Denmark Denmark "in recognition of his contribution to the treatment of diseases, especially lupus vulgaris, with concentrated light radiation, whereby he has opened a new avenue for medical science"[4]
1904 Ivan Petrovich Pavlov, Flag of Russia Russian Empire "in recognition of his work on the physiology of digestion, through which knowledge on vital aspects of the subject has been transformed and enlarged"[5]
1905 Robert Koch, Flag of German Empire German Empire "for his investigations and discoveries in relation to tuberculosis"[6]
1906 Camillo Golgi, Flag of Italy Italy;
Santiago Ramón y Cajal, Flag of Spain Spain
"in recognition of their work on the structure of the nervous system"[7]
1907 Charles Louis Alphonse Laveran, Flag of France France "in recognition of his work on the role played by protozoa in causing diseases"[8]
1908 Ilya Ilyich Mechnikov, Flag of Russia Russian Empire;
Paul Ehrlich, Flag of German Empire German Empire
"in recognition of their work on immunity"[9]
1909 Emil Theodor Kocher, Flag of Switzerland Switzerland "for his work on the physiology, pathology and surgery of the thyroid gland"[10]
1910 Albrecht Kossel, Flag of German Empire German Empire "in recognition of the contributions to our knowledge of cell chemistry made through his work on proteins, including the nucleic substances"[11]
1911 Allvar Gullstrand, Flag of Sweden Sweden "for his work on the dioptrics of the eye"[12]
1912 Alexis Carrel, Flag of France France "in recognition of his work on vascular suture and the transplantation of blood vessels and organs"[13]
1913 Charles Robert Richet, Flag of France France "in recognition of his work on anaphylaxis"[14]
1914 Robert Bárány, Flag of Austria Austria "for his work on the physiology and pathology of the vestibular apparatus"[15]
1915 [No award]
1916 [No award]
1917 [No award]
1918 [No award]
1919 Jules Bordet, Flag of Belgium Belgium "for his discoveries relating to immunity"[16]
1920 Schack August Steenberg Krogh, Flag of Denmark Denmark "for his discovery of the capillary motor regulating mechanism" (for showing that the gas exchange in the lungs is ordinary diffusion)[17]
1921 [No award]
1922* Archibald Vivian Hill, Flag of the United Kingdom United Kingdom "for his discovery relating to the production of heat in the muscle"[18]
1922* Otto Fritz Meyerhof, Flag of Weimar Republic Weimar Republic (Germany) "for his discovery of the fixed relationship between the consumption of oxygen and the metabolism of lactic acid in the muscle"[18]
1923 Frederick Grant Banting, Flag of Canada Canada;
John James Richard Macleod, Flag of the United Kingdom United Kingdom, Flag of Canada Canada?
"for the discovery of insulin"[19]
1924 Willem Einthoven, Flag of the Netherlands Netherlands "for the discovery of the mechanism of the electrocardiogram"[20]
1925 [No award]
1926 Johannes Andreas Grib Fibiger, Flag of Denmark Denmark "for his discovery of the Spiroptera carcinoma"[21]
1927 Julius Wagner-Jauregg, Flag of Austria Austria "for his discovery of the therapeutic value of malaria inoculation in the treatment of dementia paralytica"[22]
1928 Charles Jules Henri Nicolle, Flag of France France "for his work on typhus"[23]
1929* Christiaan Eijkman, Flag of the Netherlands Netherlands "for his discovery of the antineuritic vitamin"[24]
1929* Sir Frederick Gowland Hopkins, Flag of the United Kingdom United Kingdom "for his discovery of the growth-stimulating vitamins"[24]
1930 Karl Landsteiner, Flag of Austria Austria "for his discovery of human blood groups"[25]
1931 Otto Heinrich Warburg, Flag of Weimar Republic Weimar Republic (Germany) "for his discovery of the nature and mode of action of the respiratory enzyme"[26]
1932 Sir Charles Scott Sherrington, Flag of the United Kingdom United Kingdom;
Edgar Douglas Adrian, Flag of the United Kingdom United Kingdom
"for their discoveries regarding the functions of neurons"[27]
1933 Thomas Hunt Morgan, Flag of the United States United States "for his discoveries concerning the role played by the chromosome in heredity"[28]
1934 George Hoyt Whipple, Flag of the United States United States;
George Richards Minot, Flag of the United States United States;
William Parry Murphy, Flag of the United States United States
"for their discoveries concerning liver therapy in cases of anaemia"[29]
1935 Hans Spemann, Flag of Nazi Germany Nazi Germany "for his discovery of the organizer effect in embryonic development"[30]
1936 Sir Henry Hallett Dale, Flag of the United Kingdom United Kingdom;
Otto Loewi, Flag of German Empire German Empire, 1903: Flag of Austria Austria, 1946: Flag of the United States United States
"for their discoveries relating to chemical transmission of nerve impulses"[31]
1937 Albert Szent-Györgyi von Nagyrapolt, Flag of Hungary Hungary "for his discoveries in connection with the biological combustion processes, with special reference to vitamin C and the catalysis of fumaric acid"[32]
1938 Corneille Jean François Heymans, Flag of Belgium Belgium "for the discovery of the role played by the sinus and aortic mechanisms in the regulation of respiration"[33]
1939 Gerhard Domagk, Flag of Nazi Germany Nazi Germany "for the discovery of the antibacterial effects of prontosil"[34]
1940 [No award]
1941 [No award]
1942 [No award]
1943* Carl Peter Henrik Dam, Flag of Denmark Denmark;
Edward Adelbert Doisy, Flag of the United States United States
"for his discovery of vitamin K"[35]
1943* Edward Adelbert Doisy, Flag of the United States United States "for his discovery of the chemical nature of vitamin K"[35]
1944 Joseph Erlanger, Flag of the United States United States;
Herbert Spencer Gasser, Flag of the United States United States
"for their discoveries relating to the highly differentiated functions of single nerve fibres"[36]
1945 Sir Alexander Fleming, Flag of the United Kingdom United Kingdom;
Ernst Boris Chain, Flag of Nazi Germany Nazi Germany, Flag of the United Kingdom United Kingdom;
Sir Howard Walter Florey, Flag of Australia Australia, Flag of the United Kingdom United Kingdom
"for the discovery of penicillin and its curative effect in various infectious diseases"[37]
1946 Hermann Joseph Muller, Flag of the United States United States "for the discovery of the production of mutations by means of X-ray irradiation"[38]
1947* Carl Ferdinand Cori, Flag of Austria-Hungary Austria-Hungary, Flag of the United States United States;
Gerty Theresa Cori (née Radnitz), Flag of Austria-Hungary Austria-Hungary, Flag of the United States United States
"for their discovery of the course of the catalytic conversion of glycogen"[39]
1947* Bernardo Alberto Houssay, Flag of Argentina Argentina "for his discovery of the part played by the hormone of the anterior pituitary lobe in the metabolism of sugar"[39]
1948 Paul Hermann Müller, Flag of Switzerland Switzerland "for his discovery of the high efficiency of DDT as a contact poison against several arthropods"[40]
1949* Walter Rudolf Hess, Flag of Switzerland Switzerland "for his discovery of the functional organization of the interbrain as a coordinator of the activities of the internal organs"[41]
1949* Antonio Caetano De Abreu Freire Egas Moniz, Flag of Portugal Portugal "for his discovery of the therapeutic value of leucotomy in certain psychoses"[41]
1950 Edward Calvin Kendall, Flag of the United States United States;
Tadeus Reichstein, Flag of Poland Poland, Flag of Switzerland Switzerland;
Philip Showalter Hench, Flag of the United States United States
"for their discoveries relating to the hormones of the adrenal cortex, their structure and biological effects"[42]
1951 Max Theiler, Flag of South Africa South Africa "for his discoveries concerning yellow fever and how to combat it"[43]
1952 Selman Abraham Waksman, Flag of Russia Russian Empire, 1916: Flag of the United States United States "for his discovery of streptomycin, the first antibiotic effective against tuberculosis"[44]
1953* Hans Adolf Krebs, Flag of German Empire German Empire, Flag of the United Kingdom United Kingdom "for his discovery of the citric acid cycle"[45]
1953* Fritz Albert Lipmann, Flag of Weimar Republic Weimar Republic (Germany), Flag of the United States United States "for his discovery of co-enzyme A and its importance for intermediary metabolism"[45]
1954 John Franklin Enders, Flag of the United States United States;
Thomas Huckle Weller, Flag of the United States United States;
Frederick Chapman Robbins, Flag of the United States United States
"for their discovery of the ability of poliomyelitis viruses to grow in cultures of various types of tissue"[46]
1955 Axel Hugo Theodor Theorell, Flag of Sweden Sweden "for his discoveries concerning the nature and mode of action of oxidation enzymes"[47]
1956 André Frédéric Cournand, Flag of France France, 1941: Flag of the United States United States;
Werner Forssmann, Flag of West Germany West Germany;
Dickinson W. Richards, Flag of the United States United States
"for their discoveries concerning heart catheterization and pathological changes in the circulatory system"[48]
1957 Daniel Bovet, Flag of Switzerland Switzerland, Flag of Italy Italy "for his discoveries relating to synthetic compounds that inhibit the action of certain body substances, and especially their action on the vascular system and the skeletal muscles"[49]
1958* George Wells Beadle, Flag of the United States United States;
Edward Lawrie Tatum, Flag of the United States United States
"for their discovery that genes act by regulating definite chemical events"[50]
1958* Joshua Lederberg Flag of the United States United States "for his discoveries concerning genetic recombination and the organization of the genetic material of bacteria"[50]
1959 Severo Ochoa, Flag of Spain Spain, Flag of the United States United States;
Arthur Kornberg, Flag of the United States United States
"for their discovery of the mechanisms in the biological synthesis of ribonucleic acid and deoxyribonucleic acid"[51]
1960 Sir Frank Macfarlane Burnet, Flag of Australia Australia, Flag of the United Kingdom United Kingdom;
Peter Brian Medawar, Flag of Brazil Brazil, Flag of the United Kingdom United Kingdom
"for discovery of acquired immunological tolerance"[52]
1961 Georg von Békésy, Flag of Hungary Hungary "for his discoveries of the physical mechanism of stimulation within the cochlea"[53]
1962 Francis Harry Compton Crick, Flag of the United Kingdom United Kingdom;
James Dewey Watson, Flag of the United States United States
Maurice Hugh Frederick Wilkins, Flag of New Zealand New Zealand, Flag of the United Kingdom United Kingdom
"for their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material"[54]
1963 Sir John Carew Eccles, Flag of Australia Australia;
Alan Lloyd Hodgkin, Flag of the United Kingdom United Kingdom;
Andrew Fielding Huxley, Flag of the United Kingdom United Kingdom
"for their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane"[55]
1964 Konrad Bloch, Flag of German Empire German Empire, Flag of the United States United States;
Feodor Lynen, Flag of West Germany West Germany
"for their discoveries concerning the mechanism and regulation of the cholesterol and fatty acid metabolism"[56]
1965 François Jacob, Flag of France France;
André Lwoff, Flag of France France;
Jacques Monod, Flag of France France
"for their discoveries concerning genetic control of enzyme and virus synthesis"[57]
1966* Peyton Rous, Flag of the United States United States "for his discovery of tumour-inducing viruses"[58]
1966* Charles B. Huggins, Flag of Canada Canada, Flag of the United States United States "for his discoveries concerning hormonal treatment of prostatic cancer"[58]
1967 Ragnar Granit, Flag of Finland Finland, 1940: Flag of Sweden Sweden;
Haldan Keffer Hartline, Flag of the United States United States;
George Wald, Flag of the United States United States
"for their discoveries concerning the primary physiological and chemical visual processes in the eye"[59]
1968 Robert W. Holley, Flag of the United States United States;
Har Gobind Khorana, Flag of India British India (India), 1966: Flag of the United States United States;
Marshall W. Nirenberg, Flag of the United States United States
"for their interpretation of the genetic code and its function in protein synthesis"[60]
1969 Max Delbrück, Flag of West Germany West Germany, Flag of the United States United States;
Alfred Hershey, Flag of the United States United States;
Salvador E. Luria, Flag of Italy Italy
"for their discoveries concerning the replication mechanism and the genetic structure of viruses"[61]
1970 Sir Bernard Katz, Flag of Nazi Germany Nazi Germany, 1941: Flag of the United Kingdom United Kingdom;
Ulf von Euler, Flag of Sweden Sweden;
Julius Axelrod, Flag of the United States United States
"for their discoveries concerning the humoral transmittors in the nerve terminals and the mechanism for their storage, release and inactivation"[62]
1971 Earl W. Sutherland, Jr., Flag of the United States United States "for his discoveries concerning the mechanisms of the action of hormones"[63]
1972 Gerald M. Edelman, Flag of the United States United States;
Rodney R. Porter, Flag of the United Kingdom United Kingdom
"for their discoveries concerning the chemical structure of antibodies"[64]
1973 Karl von Frisch, Flag of Austria Austria;
Konrad Lorenz, Flag of Austria Austria;
Nikolaas Tinbergen, Flag of the Netherlands Netherlands
"for their discoveries concerning organization and elicitation of individual and social behaviour patterns"[65]
1974 Albert Claude, Flag of Belgium Belgium;
Christian de Duve, Flag of Belgium Belgium;
George E. Palade, Flag of Romania Romania, 1952: Flag of the United States United States
"for their discoveries concerning the structural and functional organization of the cell"[66]
1975 David Baltimore, Flag of the United States United States;
Renato Dulbecco, Flag of Italy Italy, Flag of the United States United States;
Howard Martin Temin, Flag of the United States United States
"for their discoveries concerning the interaction between tumour viruses and the genetic material of the cell"[67]
1976 Baruch S. Blumberg, Flag of the United States United States;
D. Carleton Gajdusek, Flag of the United States United States
"for their discoveries concerning new mechanisms for the origin and dissemination of infectious diseases"[68]
1977* Roger Guillemin, Flag of France France, 1965: Flag of the United States United States;
Andrew Wiktor Schally, Flag of the United States United States
"for their discoveries concerning the peptide hormone production of the brain"[69]
1977* Rosalyn Yalow, Flag of the United States United States "for the development of radioimmunoassays of peptide hormones"[69]
1978 Werner Arber, Flag of Switzerland Switzerland;
Daniel Nathans, Flag of the United States United States;
Hamilton O. Smith, Flag of the United States United States
"for the discovery of restriction enzymes and their application to problems of molecular genetics"[70]
1979 Allan M. Cormack, Flag of South Africa South Africa, 1966: Flag of the United States United States;
Godfrey N. Hounsfield, Flag of the United Kingdom United Kingdom
"for the development of computer assisted tomography"[71]
1980 Baruj Benacerraf, Flag of Venezuela Venezuela, 1943: Flag of the United States United States;
Jean Dausset, Flag of France France;
George D. Snell, Flag of the United States United States
"for their discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions"[72]
1981* Roger W. Sperry, Flag of the United States United States "for his discoveries concerning the functional specialization of the cerebral hemispheres"[73]
1981* David H. Hubel, Flag of the United States United States;
Torsten N. Wiesel, Flag of Sweden Sweden
"for their discoveries concerning information processing in the visual system"[73]
1982 Sune Bergström, Flag of Sweden Sweden;
Bengt I. Samuelsson, Flag of Sweden Sweden;
John R. Vane, Flag of the United Kingdom United Kingdom
"for their discoveries concerning prostaglandins and related biologically active substances"[74]
1983 Barbara McClintock, Flag of the United States United States "for her discovery of mobile genetic elements"[75]
1984 Niels K. Jerne, Flag of Denmark Denmark;
Georges J.F. Köhler, Flag of West Germany West Germany;
César Milstein, Flag of Argentina Argentina, Flag of the United Kingdom United Kingdom
"for theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies"[76]
1985 Michael S. Brown, Flag of the United States United States;
Joseph L. Goldstein, Flag of the United States United States
"for their discoveries concerning the regulation of cholesterol metabolism"[77]
1986 Stanley Cohen, Flag of the United States United States;
Rita Levi-Montalcini, Flag of Italy Italy, Flag of the United States United States
"for their discoveries of growth factors"[78]
1987 Susumu Tonegawa, Flag of Japan Japan "for his discovery of the genetic principle for generation of antibody diversity"[79]
1988 Sir James W. Black, Flag of the United Kingdom United Kingdom;
Gertrude B. Elion, Flag of the United States United States;
George H. Hitchings, Flag of the United States United States
"for their discoveries of important principles for drug treatment"[80]
1989 J. Michael Bishop, Flag of the United States United States;
Harold E. Varmus, Flag of the United States United States
"for their discovery of the cellular origin of retroviral oncogenes"[81]
1990 Joseph E. Murray, Flag of the United States United States;
E. Donnall Thomas, Flag of the United States United States
"for their discoveries concerning organ and cell transplantation in the treatment of human disease"[82]
1991 Erwin Neher, Flag of Germany Germany;
Bert Sakmann, Flag of Germany Germany
"for their discoveries concerning the function of single ion channels in cells"[83]
1992 Edmond H. Fischer, Flag of Switzerland Switzerland, Flag of the United States United States;
Edwin G. Krebs, Flag of the United States United States
"for their discoveries concerning reversible protein phosphorylation as a biological regulatory mechanism"[84]
1993 Richard J. Roberts, Flag of the United Kingdom United Kingdom;
Phillip A. Sharp, Flag of the United States United States
"for their discoveries of split genes"[85]
1994 Alfred G. Gilman, Flag of the United States United States;
Martin Rodbell, Flag of the United States United States
"for their discovery of G-proteins and the role of these proteins in signal transduction in cells"[86]
1995 Edward B. Lewis, Flag of the United States United States;
Christiane Nüsslein-Volhard, Flag of Germany Germany;
Eric F. Wieschaus, Flag of the United States United States
"for their discoveries concerning the genetic control of early embryonic development"[87]
1996 Peter C. Doherty, Flag of Australia Australia;
Rolf M. Zinkernagel, Flag of Switzerland Switzerland
"for their discoveries concerning the specificity of the cell mediated immune defence"[88]
1997 Stanley B. Prusiner, Flag of the United States United States "for his discovery of Prions - a new biological principle of infection"[89]
1998 Robert F. Furchgott, Flag of the United States United States;
Louis J. Ignarro, Flag of the United States United States;
Ferid Murad, Flag of the United States United States
"for their discoveries concerning nitric oxide as a signalling molecule in the cardiovascular system"[90]
1999 Günter Blobel, Flag of West Germany West Germany, 1987: Flag of the United States United States "for the discovery that proteins have intrinsic signals that govern their transport and localization in the cell"[91]
2000 Arvid Carlsson, Flag of Sweden Sweden;
Paul Greengard, Flag of the United States United States;
Eric R. Kandel, Flag of the United States United States
"for their discoveries concerning signal transduction in the nervous system"[92]
2001 Leland H. Hartwell, Flag of the United States United States;
R. Timothy (Tim) Hunt, Flag of the United Kingdom United Kingdom;
Sir Paul M. Nurse, Flag of the United Kingdom United Kingdom
"for their discoveries of key regulators of the cell cycle"[93]
2002 Sydney Brenner, Flag of South Africa South Africa, Flag of the United Kingdom United Kingdom, 2003: Flag of Singapore Singapore[94] (honorary);
H. Robert Horvitz, Flag of the United States United States;
John E. Sulston, Flag of the United Kingdom United Kingdom
"for their discoveries concerning 'genetic regulation of organ development and programmed cell death'"[95]
2003 Paul Lauterbur, Flag of the United States United States;
Sir Peter Mansfield, Flag of the United Kingdom United Kingdom
"for their discoveries concerning magnetic resonance imaging"[96]
2004 Richard Axel, Flag of the United States United States;
Linda B. Buck, Flag of the United States United States
"for their discoveries of odorant receptors and the organization of the olfactory system"[97]
2005 Barry J. Marshall, Flag of Australia Australia;
J. Robin Warren, Flag of Australia Australia
"for their discovery of the bacterium Helicobacter pylori and its role in gastritis and peptic ulcer disease"[98]
2006 Andrew Z. Fire, Flag of the United States United States;
Craig C. Mello, Flag of the United States United States
"for their discovery of RNA interference - gene silencing by double-stranded RNA"[99]

* Years with multiple reasons for a Nobel prize.

[edit] References

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